In ancient Greece, having a second toe longer than the big toe was considered a standard of beauty.
This condition, today known as Morton's toe or "Greek Foot", is depicted in many famous statues, such as Michelangelo's David and the Statue of Liberty.
It is a commonly-inherited but inefficient foot shape. It's not the actual toe bones that are longer, though; rather, it's an issue with the metatarsal bones they are attached to.